Croc 2 features gameplay similar to its predecessor; the player controls Croc through various open-ended levels in order to complete various missions. The game's levels are split across 4 Gobbo "villages" and are accessed through an open HUB world.[3] The missions involve helping other Gobbos with a certain task, such as rescuing another Gobbo who is trapped within a steel cage and chasing a Dantini through a stage in order to retrieve a stolen sandwich. After completing a certain number of levels in a village, a boss level is opened up, which allows access to the next village when completed. There's also levels requiring players to control vehicles such as; a go-kart, a speedboat, a hang glider, a hot air balloon and a giant snowball among others.[3][4]
Levels contain various collectible items, including 100 crystals and 5 multicolored crystals scattered throughout the stage. The colored gems are hidden in different places throughout the stage, and require completing a platform challenge or completing a puzzle in order to be retrieved. Finding all 5 colored gems makes a golden trophy appear at the end of the level that is collected by traversing through a small platforming challenge. Collecting every trophy in a village allows access to an extra level that can be completed in order to collect a Jigsaw puzzle piece; collecting these pieces is required in order to access the game's fifth and final world consisting of remixed level segments. Several items can be purchased at the HUB worlds from Swap Meet Pete, an anthropomorphic cat, some of which are needed to access certain areas and secrets within the game. Among these items are heart pots, which lengthen Croc's maximum life count, Gummi Savers Jumps, which can be used as a trampoline in order to reach certain ledges, and the Clockwork Gobbo, a small wind-up robot that can be controlled to collect items by being used on a certain pedestal.